Entry BACSU00903 (PERR_BACSU)

BBacillus subtilis (strain 168)


OMA Group (Proteins all orthologous to each other) ?

OMA Group 774194 ?

ferric uptake regulation protein


This group has 623 members: B623

Fingerprint: CVAVDDK

L
O
K
A
C
T
H
O
A
R
A
C
I
S
3
C
A
L
L
D
A
E
R
P
E
P
Y
R
O
C
A
C
I
H
W
M
E
T
C
R
M
E
T
S
5
S
U
L
A
C
S
U
L
I
H
S
U
L
I
D
S
U
L
I
L
S
U
L
I
M
S
U
L
I
A
S
U
L
I
K
S
U
L
I
R
S
U
L
I
Y
S
U
L
I
N
S
U
L
S
9
S
U
L
S
O
S
U
L
T
O
C
A
L
M
Q
P
Y
R
A
E
P
Y
R
A
R
P
Y
R
C
J
P
Y
R
I
L
P
Y
R
N
V
P
Y
R
O
T
T
H
E
T
K
T
H
E
U
7
V
U
L
D
I
V
U
L
M
7
A
R
C
F
U
F
E
R
P
A
M
E
T
H
6
M
E
T
S
G
M
E
T
H
D
M
E
T
A
C
M
E
T
B
F
P
Y
R
F
U
P
Y
R
S
N
T
H
E
B
M
T
H
E
L
N
T
H
E
A
C
T
H
E
V
O
A
C
I
C
5
K
O
R
V
E
G
R
A
M
M
G
R
A
T
M
T
E
R
R
K
T
E
R
S
S
C
H
L
T
F
S
O
L
U
E
A
C
I
F
D
A
C
I
C
1
T
H
E
B
D
A
R
C
H
D
M
O
B
C
V
B
I
F
A
A
B
I
F
A
R
B
I
F
A
0
B
I
F
A
B
B
I
F
L
B
B
I
F
A
S
B
I
F
A
V
B
I
F
A
P
B
I
F
B
P
B
I
F
B
S
B
I
F
B
A
B
I
F
D
B
B
I
F
L
D
B
I
F
L
O
B
I
F
L
1
B
I
F
L
S
B
I
F
L
2
B
I
F
L
M
B
I
F
L
J
G
A
R
V
4
G
A
R
V
3
G
A
R
V
H
C
A
T
A
D
A
T
O
P
D
O
L
S
U
V
C
O
R
G
P
C
R
Y
C
D
E
G
G
L
E
S
L
A
H
D
C
O
R
A
7
C
O
R
D
2
C
O
R
D
3
C
O
R
D
7
C
O
R
D
I
C
O
R
D
D
C
O
R
D
H
C
O
R
D
J
C
O
R
D
K
C
O
R
D
L
C
O
R
D
W
C
O
R
D
V
C
O
R
E
F
C
O
R
G
L
C
O
R
G
K
C
O
R
G
B
C
O
R
J
K
C
O
R
K
4
C
O
R
P
1
C
O
R
P
2
C
O
R
P
F
C
O
R
P
9
C
O
R
R
G
C
O
R
U
B
C
O
R
U
7
C
O
R
V
D
G
O
R
B
4
G
O
R
P
V
H
O
Y
S
D
M
Y
C
A
S
M
Y
C
A
1
M
Y
C
P
A
M
Y
C
I
A
M
Y
C
L
B
M
Y
C
L
E
M
Y
C
M
M
M
Y
C
S
J
M
Y
C
S
K
M
Y
C
S
S
M
Y
C
A
0
M
Y
C
B
O
M
Y
C
B
P
M
Y
C
B
T
M
Y
C
C
P
M
Y
C
T
A
M
Y
C
T
U
M
Y
C
T
C
M
Y
C
T
D
M
Y
C
T
F
M
Y
C
T
K
M
Y
C
U
A
M
Y
C
A
9
M
Y
C
S
D
M
Y
C
R
N
M
Y
C
S
2
M
Y
C
V
P
M
Y
C
C
N
M
Y
C
S
R
M
Y
C
G
I
N
O
C
C
G
N
O
C
F
A
R
H
O
E
4
R
H
O
H
1
R
H
O
J
R
R
H
O
O
B
S
E
G
R
D
T
S
U
P
D
F
R
A
A
A
F
R
A
C
C
F
R
A
I
E
F
R
A
S
N
F
R
A
D
G
B
L
A
S
D
G
E
O
O
G
M
O
D
M
B
S
T
A
N
L
K
I
N
R
D
B
E
U
C
1
B
R
E
L
N
C
E
L
F
A
C
E
L
F
N
C
E
L
G
A
B
R
A
F
D
K
Y
T
S
D
J
O
N
D
D
C
L
A
M
3
C
L
A
M
S
L
E
I
X
X
M
I
C
T
S
A
R
T
S
2
G
L
U
A
R
K
O
C
R
D
M
I
C
L
C
P
A
E
A
T
P
S
E
C
P
P
S
E
P
M
R
E
N
S
M
R
O
T
D
C
R
O
T
M
D
I
S
O
V
2
X
Y
L
C
X
S
A
N
K
S
A
C
T
M
4
A
C
T
S
5
M
I
C
A
I
M
I
C
S
L
S
A
L
A
I
S
A
L
T
O
V
E
R
M
A
N
A
K
M
Y
K
R
I
F
D
N
O
C
S
J
A
C
I
A
4
C
U
T
A
K
P
R
O
A
S
M
I
C
P
N
P
R
O
F
C
P
S
E
P
Q
A
C
T
M
D
A
M
Y
M
S
A
M
Y
M
U
P
S
E
U
X
S
A
C
V
D
S
A
C
E
N
S
A
C
E
S
R
U
B
X
D
K
I
T
S
K
S
T
R
A
W
S
T
R
B
B
S
T
R
E
N
S
T
R
C
O
S
T
R
G
G
S
T
R
H
J
S
T
R
F
A
S
T
R
S
W
S
T
R
V
P
N
O
C
A
A
N
O
C
D
D
T
H
E
F
Y
S
T
R
R
D
T
H
E
C
D
C
O
N
W
I
A
Q
U
A
E
H
Y
D
T
T
H
Y
D
S
0
T
H
E
A
H
P
E
R
M
H
S
U
L
A
A
S
U
L
S
Y
D
E
S
T
D
T
H
E
A
1
R
H
O
M
4
S
A
L
R
D
S
A
L
R
M
B
A
C
F
6
B
A
C
F
N
B
A
C
F
R
B
A
C
T
6
B
A
C
S
H
B
A
C
T
N
B
A
C
V
8
O
D
O
S
D
P
A
L
P
W
P
O
R
A
D
P
O
R
E
A
P
O
R
G
3
P
O
R
G
I
P
R
E
D
F
P
R
E
I
7
P
R
E
M
B
P
R
E
R
2
P
A
R
D
8
T
A
N
F
A
C
H
I
P
D
N
I
A
K
G
B
E
R
L
S
B
E
L
B
D
C
Y
C
M
S
E
C
H
V
K
C
Y
T
H
3
D
Y
A
F
D
E
M
T
O
G
L
E
A
B
4
R
U
N
S
L
S
P
I
L
D
M
A
R
T
H
F
L
U
T
R
O
W
E
H
D
A
E
Q
S
U
C
A
P
C
C
C
A
P
O
D
C
E
L
A
D
C
E
L
L
C
C
R
O
A
H
D
O
K
S
4
F
L
A
B
F
F
L
A
C
A
F
L
A
I
G
F
L
A
J
1
F
L
A
P
J
G
R
A
F
K
L
A
C
S
5
L
E
E
B
M
M
A
R
S
H
M
U
R
R
D
N
O
N
D
D
O
R
N
R
L
P
S
Y
T
T
R
I
E
A
R
R
O
B
B
H
W
E
E
V
C
Z
O
B
G
A
Z
U
N
P
S
F
L
A
B
3
H
A
L
H
1
S
A
P
G
L
P
E
D
H
D
P
S
E
S
L
S
O
L
C
M
S
P
H
S
2
C
A
L
E
A
P
A
R
U
W
W
A
D
C
W
C
H
L
P
8
C
H
L
T
E
C
H
L
C
H
C
H
L
L
2
C
H
L
P
B
C
H
L
P
D
C
H
L
P
M
C
H
L
T
3
C
H
L
L
7
P
E
L
P
B
P
R
O
A
2
A
N
A
T
U
C
A
L
A
S
C
H
L
A
D
C
H
L
S
Y
C
H
L
A
A
R
O
S
C
S
R
O
S
S
1
H
E
R
A
2
D
E
H
M
B
D
E
H
M
1
D
E
H
M
C
D
E
H
M
G
D
E
H
M
V
D
E
H
L
B
S
P
H
T
D
T
H
E
R
P
D
E
S
I
S
G
L
O
V
I
A
N
A
C
C
N
O
S
P
7
N
O
S
S
7
N
O
S
S
1
A
N
A
V
T
N
O
S
A
0
H
A
L
P
7
C
Y
A
A
P
C
Y
A
S
C
M
I
C
A
N
C
Y
A
A
5
C
Y
A
P
7
C
Y
A
P
4
C
Y
A
P
2
C
Y
A
P
8
C
Y
A
P
0
A
R
T
P
N
T
R
I
E
I
L
Y
N
S
P
S
T
A
C
7
A
C
A
M
1
C
H
A
P
6
S
Y
N
Y
3
P
R
O
M
S
P
R
O
M
4
P
R
O
M
2
P
R
O
M
0
P
R
O
M
3
P
R
O
M
9
P
R
O
M
M
P
R
O
M
5
P
R
O
M
1
P
R
O
M
T
P
R
O
M
A
P
R
O
M
P
C
Y
A
G
P
S
Y
N
P
6
S
Y
N
P
2
S
Y
N
S
7
S
Y
N
S
3
S
Y
N
S
C
S
Y
N
S
9
S
Y
N
J
B
S
Y
N
J
A
S
Y
N
R
3
S
Y
N
P
W
S
Y
N
P
X
T
H
E
E
B
C
A
L
N
Y
D
E
F
D
S
D
E
N
A
2
F
L
E
S
M
D
E
I
D
V
D
E
I
G
D
D
E
I
G
I
D
E
I
M
L
D
E
I
P
D
D
E
I
P
M
D
E
I
R
A
T
R
U
R
R
M
A
R
H
T
M
E
I
R
D
M
E
I
S
D
O
C
E
P
5
T
H
E
T
2
T
H
E
T
8
T
H
E
T
G
D
I
C
T
6
D
I
C
T
D
E
L
U
M
P
F
I
B
S
S
A
L
I
A
T
A
L
I
A
D
K
Y
R
T
2
A
M
P
X
N
A
N
O
F
W
B
A
C
A
3
B
A
C
A
S
B
A
C
A
N
B
A
C
A
A
B
A
C
A
C
B
A
C
A
1
B
A
C
C
J
B
A
C
C
3
B
A
C
C
7
B
A
C
C
0
B
A
C
C
1
B
A
C
C
R
B
A
C
C
4
B
A
C
C
2
B
A
C
C
Q
B
A
C
C
Z
B
A
C
A
I
B
A
C
S
K
B
A
C
C
6
B
A
C
C
N
B
A
C
H
D
B
A
C
L
D
B
A
C
M
Q
B
A
C
M
D
B
A
C
W
K
B
A
C
P
E
B
A
C
P
2
B
A
C
S
T
B
A
C
P
Z
B
A
C
P
T
B
A
C
S
U
B
A
C
A
H
B
A
C
T
0
B
A
C
H
K
B
A
C
T
1
B
A
C
V
Z
G
E
O
K
A
G
E
O
S
W
G
E
O
S
0
G
E
O
S
Y
G
E
O
T
N
L
Y
S
S
C
O
C
E
I
H
G
E
O
T
C
E
X
I
A
B
E
X
I
S
2
E
X
I
S
A
L
I
S
I
N
L
I
S
I
P
L
I
S
M
2
L
I
S
M
4
L
I
S
M
H
L
I
S
M
L
L
I
S
M
M
L
I
S
M
C
L
I
S
M
F
L
I
S
M
O
L
I
S
S
S
L
I
S
W
6
B
R
E
B
N
G
E
O
S
4
P
A
E
M
K
P
A
E
P
6
P
A
E
P
S
P
A
E
S
J
P
A
E
T
H
T
H
E
C
K
S
O
L
S
S
B
A
C
I
E
M
A
C
C
J
S
T
A
A
4
S
T
A
A
B
S
T
A
A
C
S
T
A
A
G
S
T
A
A
D
S
T
A
A
2
S
T
A
A
9
S
T
A
A
K
S
T
A
A
J
S
T
A
A
5
S
T
A
A
R
S
T
A
A
S
S
T
A
A
W
S
T
A
A
1
S
T
A
A
M
S
T
A
A
N
S
T
A
A
8
S
T
A
A
E
S
T
A
A
H
S
T
A
A
0
S
T
A
A
3
S
T
A
A
T
S
T
A
A
F
S
T
A
C
T
S
T
A
E
S
S
T
A
E
Q
S
T
A
H
J
S
T
A
H
Y
S
T
A
L
H
S
T
A
P
E
S
T
A
P
H
S
T
A
S
1
A
E
R
U
A
C
A
R
S
1
E
N
T
F
6
E
N
T
F
O
E
N
T
F
A
E
N
T
F
U
M
E
L
P
T
M
E
L
P
D
T
E
T
H
N
L
A
C
B
A
L
A
C
B
N
L
A
C
C
D
L
A
C
C
B
L
A
C
C
C
L
A
C
C
Z
L
A
C
F
3
L
A
C
P
3
L
A
C
P
L
L
A
C
P
J
L
A
C
P
S
L
A
C
R
S
L
A
C
R
D
L
A
C
R
J
L
A
C
R
G
L
A
C
R
L
L
A
C
R
R
L
A
C
S
S
L
A
C
S
1
L
A
C
S
M
P
E
D
C
P
P
E
D
P
A
L
E
U
C
J
L
E
U
C
K
L
E
U
G
J
L
E
U
G
G
L
E
U
M
M
L
E
U
S
2
W
E
I
K
K
W
E
I
V
I
L
A
C
L
M
L
A
C
L
N
L
A
C
L
S
L
A
C
L
V
L
A
C
L
A
L
A
C
L
K
S
T
R
A
3
S
T
R
A
1
S
T
R
A
5
S
T
R
A
2
S
T
R
E
D
S
T
R
D
G
S
T
R
E
4
S
T
R
E
C
S
T
R
S
7
S
T
R
E
M
S
T
R
G
1
S
T
R
G
2
S
T
R
G
3
S
T
R
G
C
S
T
R
I
C
S
T
R
I
J
S
T
R
M
D
S
T
R
M
U
S
T
R
M
N
S
T
R
P
W
S
T
R
P
X
S
T
R
P
1
S
T
R
P
B
S
T
R
P
C
S
T
R
P
8
S
T
R
P
D
S
T
R
P
M
S
T
R
P
3
S
T
R
P
Q
S
T
R
P
F
S
T
R
P
Z
S
T
R
P
G
S
T
R
P
6
S
T
R
E
5
S
T
R
E
H
S
T
R
E
8
S
T
R
S
V
S
T
R
S
Y
S
T
R
S
2
S
T
R
S
4
S
T
R
G
Z
S
T
R
E
J
S
T
R
S
E
S
T
R
S
X
S
T
R
T
2
S
T
R
T
D
S
T
R
T
1
S
T
R
T
N
S
T
R
U
0
A
L
K
M
Q
A
L
K
O
O
A
R
T
S
S
C
L
O
A
B
C
L
O
A
E
C
L
O
B
8
C
L
O
B
0
C
L
O
B
1
C
L
O
B
H
C
L
O
B
J
C
L
O
B
M
C
L
O
B
6
C
L
O
B
B
C
L
O
B
K
C
L
O
B
A
C
L
O
B
2
C
L
O
B
L
C
L
O
C
7
C
L
O
K
5
C
L
O
L
D
C
L
O
N
N
C
L
O
P
E
C
L
O
P
1
C
L
O
P
S
C
L
O
S
S
C
L
O
T
E
S
U
L
A
D
S
U
L
A
T
T
H
E
M
7
A
C
E
W
D
E
U
B
L
K
E
U
B
E
2
H
E
L
M
I
B
U
T
P
B
C
E
L
L
D
C
L
O
S
W
L
A
C
P
7
R
O
S
H
A
A
G
A
R
V
O
S
C
V
S
D
E
S
D
L
D
E
S
H
D
D
E
S
H
Y
D
E
S
A
S
D
E
S
K
7
D
E
S
A
J
D
E
S
M
D
D
E
S
O
D
D
E
S
C
C
D
E
S
R
M
D
E
S
R
L
P
E
L
T
S
S
Y
N
G
F
T
H
E
P
J
A
C
E
S
D
P
E
P
D
6
P
E
P
D
R
F
I
L
A
D
E
T
H
H
Y
M
A
G
I
U
C
L
O
T
H
C
L
O
T
L
C
L
O
C
E
C
L
O
C
D
R
U
M
A
7
S
Y
M
T
H
S
Y
N
W
W
S
Y
N
L
T
H
A
L
H
G
H
A
L
P
G
H
A
L
O
H
A
C
E
A
Z
H
A
L
H
C
N
A
T
T
J
C
A
L
S
4
C
A
R
H
Z
M
O
O
T
A
T
E
P
A
E
T
H
E
P
S
T
H
E
I
A
T
H
E
M
3
T
H
E
P
3
T
H
E
S
X
T
H
E
P
X
C
A
L
B
D
C
A
L
H
1
C
A
L
K
I
C
A
L
K
2
C
A
L
O
O
C
A
L
O
W
C
A
L
S
8
T
H
E
S
W
T
H
E
T
C
T
H
E
X
L
T
H
E
O
J
M
A
H
A
5
E
R
Y
R
F
A
C
I
F
V
A
C
I
I
R
S
E
L
R
L
S
E
L
S
3
V
E
I
P
T
C
L
O
A
9
A
N
A
P
D
F
I
N
M
2
F
U
S
N
N
I
L
Y
P
C
L
E
P
B
D
S
E
B
T
E
S
T
R
M
9
G
E
M
A
T
I
G
N
A
J
M
E
L
R
P
L
E
P
F
C
L
E
P
F
M
N
I
T
M
O
T
H
E
Y
D
P
H
Y
M
F
I
S
O
P
I
S
I
N
A
D
P
I
R
S
D
P
L
A
L
2
R
H
O
B
A
R
U
B
B
R
A
C
I
C
S
A
C
I
F
2
A
C
I
F
5
A
S
T
E
C
B
R
E
S
C
C
A
U
S
T
C
A
U
S
K
C
A
U
V
C
C
A
U
V
N
P
H
E
Z
H
M
A
G
M
M
P
A
R
B
H
P
E
L
S
M
P
E
L
U
B
B
A
R
B
K
B
A
R
C
7
B
A
R
G
A
B
A
R
H
E
B
A
R
Q
U
B
A
R
T
1
B
A
R
V
W
B
E
I
I
9
M
E
T
S
B
B
R
A
D
U
B
R
A
S
B
B
R
A
S
O
N
I
T
H
X
N
I
T
W
N
O
L
I
C
O
O
L
I
C
M
R
H
O
P
A
R
H
O
P
5
R
H
O
P
B
R
H
O
P
S
R
H
O
P
X
R
H
O
P
2
R
H
O
P
T
B
R
U
A
2
B
R
U
A
1
B
R
U
A
B
B
R
U
C
2
B
R
U
M
5
B
R
U
M
E
B
R
U
M
B
B
R
U
M
C
B
R
U
O
2
B
R
U
S
U
B
R
U
S
I
O
C
H
A
4
B
L
A
V
I
H
Y
P
D
A
H
Y
P
S
M
P
E
L
H
B
R
H
O
V
T
M
E
T
N
O
M
E
T
R
J
M
E
T
S
4
M
E
T
E
A
M
E
T
C
4
M
E
T
E
D
M
E
T
E
P
M
E
T
P
B
M
E
T
S
Z
C
H
E
S
B
M
E
S
A
W
M
E
S
C
W
R
H
I
L
O
A
G
R
S
H
A
G
R
F
C
A
G
R
R
K
A
G
R
V
S
R
H
I
E
C
R
H
I
E
6
R
H
I
L
S
R
H
I
L
W
R
H
I
L
3
R
H
I
F
H
S
I
N
F
N
R
H
I
M
E
S
I
N
M
K
S
I
N
M
B
S
I
N
M
M
S
I
N
M
W
P
A
R
L
1
A
Z
O
C
5
S
T
A
N
D
X
A
N
P
2
H
I
R
B
I
H
Y
P
N
A
M
A
R
M
M
D
I
N
S
H
J
A
N
S
C
K
E
T
V
W
K
E
T
V
Y
O
C
E
G
H
P
A
R
D
P
P
H
A
I
B
P
S
E
U
V
R
H
O
C
B
R
H
O
S
4
R
H
O
S
5
R
H
O
S
1
R
H
O
S
K
R
O
S
D
O
R
O
S
L
O
R
U
E
P
O
R
U
E
S
T
A
C
E
P
3
A
C
I
C
J
A
C
I
M
A
G
L
U
D
A
G
L
U
O
X
G
R
A
B
C
K
O
M
M
N
A
Z
O
L
4
M
A
G
S
A
R
H
O
C
S
R
H
O
R
T
T
I
S
M
K
M
I
D
M
I
E
R
Y
L
H
N
O
V
A
D
S
P
H
J
U
S
P
H
W
W
S
P
H
A
L
Z
Y
M
M
A
Z
Y
M
M
O
Z
Y
M
M
N
Z
Y
M
M
T
M
I
C
A
A
P
O
L
G
S
P
U
N
M
I
A
C
H
X
A
A
D
V
K
W
B
O
R
A
1
B
O
R
B
R
B
O
R
B
M
B
O
R
P
A
B
O
R
P
1
B
O
R
P
C
B
O
R
P
E
B
O
R
P
D
P
U
S
S
T
T
A
Y
E
M
B
U
R
C
M
B
U
R
A
4
B
U
R
C
J
B
U
R
C
A
B
U
R
C
H
B
U
R
C
C
B
U
R
L
3
B
U
R
M
1
B
U
R
V
G
B
U
R
G
S
B
U
R
G
B
B
U
R
M
A
B
U
R
M
9
B
U
R
M
7
B
U
R
M
S
B
U
R
P
0
B
U
R
P
1
B
U
R
P
6
B
U
R
P
S
B
U
R
S
C
B
U
R
S
G
B
U
R
T
A
C
U
P
M
C
C
U
P
N
H
C
U
P
N
N
C
U
P
N
J
C
U
P
T
R
P
A
R
P
8
P
A
R
P
J
P
A
R
R
H
P
A
R
X
L
P
O
L
A
Q
P
O
L
N
S
R
A
L
P
1
R
A
L
P
J
R
A
L
S
L
R
A
L
S
O
R
A
L
S
8
A
C
I
A
P
A
C
I
A
C
A
C
I
E
T
A
C
I
S
J
A
L
I
D
K
C
O
M
T
2
D
E
L
A
S
D
E
L
S
C
P
O
L
N
A
P
O
L
S
J
R
A
M
T
T
R
H
O
F
T
V
A
R
P
E
V
A
R
P
S
V
E
R
E
I
C
O
L
F
T
H
E
R
S
S
H
E
R
A
R
J
A
N
M
A
L
E
P
C
P
M
E
T
P
P
R
U
B
G
I
T
H
I
K
1
C
H
R
V
O
L
A
R
H
H
P
S
E
U
L
N
E
I
G
1
N
E
I
G
2
N
E
I
M
L
N
E
I
M
A
N
E
I
M
W
N
E
I
M
B
N
E
I
M
7
N
E
I
M
H
N
E
I
M
G
N
E
I
M
M
N
E
I
M
P
N
E
I
M
O
N
E
I
M
N
N
E
I
M
0
N
E
I
M
8
N
E
I
M
F
G
A
L
C
S
S
I
D
L
E
M
E
T
F
K
M
E
T
M
L
M
E
T
G
S
M
E
T
S
6
N
I
T
E
U
N
I
T
E
C
N
I
T
S
I
N
I
T
M
U
T
H
I
D
A
D
E
C
A
R
A
R
O
A
E
D
E
C
S
P
A
Z
O
S
B
T
H
A
S
P
A
C
C
P
U
A
E
R
H
H
A
E
R
S
4
A
E
R
V
B
O
C
E
S
G
T
O
L
A
T
A
L
T
M
B
A
L
T
M
S
A
L
T
M
E
A
L
T
M
D
A
L
T
N
A
M
A
R
A
H
M
A
R
H
V
C
O
L
P
3
F
E
R
B
D
I
D
I
L
O
P
S
E
A
6
P
S
E
H
T
P
S
E
U
9
P
S
Y
I
N
S
H
E
A
M
S
H
E
B
5
S
H
E
B
8
S
H
E
B
9
S
H
E
B
2
S
H
E
B
6
S
H
E
D
O
S
H
E
F
N
S
H
E
H
H
S
H
E
L
P
S
H
E
O
N
S
H
E
P
A
S
H
E
P
W
S
H
E
P
2
S
H
E
P
C
S
H
E
S
H
S
H
E
S
A
S
H
E
S
M
S
H
E
S
R
S
H
E
S
W
S
H
E
V
D
S
H
E
W
M
C
A
R
H
6
D
I
C
N
V
C
E
L
J
U
S
A
C
D
2
S
I
M
A
S
T
E
R
T
T
A
L
L
V
D
N
I
T
H
N
N
I
T
O
C
N
I
T
W
C
T
H
I
V
6
A
L
K
E
H
H
A
L
H
L
T
H
I
S
K
T
H
I
S
H
H
A
L
N
C
C
I
T
K
8
C
I
T
R
I
C
R
O
S
8
C
R
O
T
Z
E
N
T
A
L
E
N
T
C
C
E
N
T
3
8
E
C
O
L
X
E
C
O
C
1
E
C
O
C
2
E
C
O
5
5
E
C
O
D
1
E
C
O
K
O
E
C
O
L
W
E
C
O
B
B
E
C
O
B
D
E
C
O
L
I
E
C
O
D
H
E
C
O
B
W
E
C
O
S
E
E
C
O
S
M
E
C
O
U
M
E
C
O
U
T
E
C
O
B
R
E
C
O
L
C
E
C
O
1
0
E
C
O
1
E
E
C
O
1
A
E
C
O
2
7
E
C
O
2
4
E
C
O
S
5
E
C
O
5
7
E
C
O
5
E
E
C
O
5
T
E
C
O
L
U
E
C
O
K
I
E
C
O
K
1
E
C
O
2
6
E
C
O
4
4
E
C
O
4
5
E
C
O
C
B
E
C
O
L
6
E
C
O
L
5
E
C
O
H
1
E
C
O
7
I
E
C
O
8
A
E
C
O
8
1
E
C
O
8
N
E
C
O
H
S
E
C
O
A
B
E
S
C
F
3
K
L
E
A
K
K
L
E
O
K
K
L
E
P
3
K
L
E
P
7
K
L
E
P
H
K
L
E
V
T
E
N
T
L
S
S
A
L
B
C
S
A
L
D
C
S
A
L
A
4
S
A
L
C
H
S
A
L
P
S
S
A
L
H
S
S
A
L
P
B
S
A
L
P
C
S
A
L
S
V
S
A
L
E
P
S
A
L
G
2
S
A
L
N
S
S
A
L
P
K
S
A
L
P
A
S
A
L
T
I
S
A
L
T
4
S
A
L
T
D
S
A
L
T
Y
S
A
L
T
1
S
A
L
T
S
S
A
L
A
R
S
H
I
B
3
S
H
I
B
S
S
H
I
D
S
S
H
I
F
L
S
H
I
F
8
S
H
I
F
2
S
H
I
S
S
S
H
I
B
C
B
L
O
F
L
B
L
O
P
B
B
L
O
V
B
H
A
M
D
5
E
N
T
B
F
E
R
W
A
E
E
R
W
A
C
E
R
W
B
E
E
R
W
P
6
E
R
W
P
E
E
R
W
S
E
E
R
W
T
9
P
A
N
A
A
P
A
N
A
M
P
A
N
S
A
P
A
N
V
C
E
D
W
I
9
E
D
W
T
E
E
D
W
T
F
P
H
O
A
A
P
H
O
L
L
P
R
O
M
H
P
R
O
R
D
P
R
O
S
M
X
E
N
B
S
X
E
N
N
A
D
I
C
C
1
D
I
C
D
3
D
I
C
P
7
D
I
C
Z
5
P
E
C
A
S
P
E
C
C
P
P
E
C
S
S
P
E
C
P
W
S
O
D
G
M
R
A
H
A
C
R
A
H
S
Y
S
E
R
F
O
S
E
R
S
A
S
E
R
P
5
Y
E
R
E
8
Y
E
R
E
1
Y
E
R
E
3
Y
E
R
P
E
Y
E
R
P
D
Y
E
R
P
1
Y
E
R
P
P
Y
E
R
P
Z
Y
E
R
P
G
Y
E
R
P
A
Y
E
R
P
N
Y
E
R
P
H
Y
E
R
P
S
Y
E
R
P
B
Y
E
R
P
3
Y
E
R
P
Y
Y
E
R
R
U
C
O
X
B
2
C
O
X
B
1
C
O
X
B
N
C
O
X
B
R
C
O
X
B
U
L
E
G
L
N
L
E
G
P
C
L
E
G
P
L
L
E
G
P
A
L
E
G
P
2
L
E
G
P
H
T
A
T
M
I
M
E
T
C
A
M
E
T
A
A
M
E
T
A
2
M
E
T
M
M
A
L
C
B
S
A
L
C
D
B
H
A
H
C
H
C
H
R
S
D
H
A
L
E
D
K
A
N
K
D
M
A
R
M
1
M
A
R
M
S
N
E
P
C
E
A
C
T
P
J
A
C
T
P
2
A
C
T
P
7
A
C
T
S
Z
A
G
G
A
D
A
G
G
A
N
M
A
N
S
M
G
A
L
A
U
H
A
E
P
S
H
A
E
D
U
H
A
E
I
1
H
A
E
I
8
H
A
E
I
N
H
A
E
I
E
H
A
E
I
G
H
A
E
I
2
H
A
E
I
6
H
A
E
P
3
H
A
E
S
1
H
I
S
S
2
P
A
S
M
H
P
A
S
M
U
A
C
I
A
D
A
C
I
B
1
A
C
I
B
5
A
C
I
B
3
A
C
I
B
C
A
C
I
B
T
A
C
I
B
Y
A
C
I
B
S
A
C
I
B
D
A
C
I
C
P
A
C
I
S
D
M
O
R
C
R
M
O
R
N
O
P
S
Y
A
2
P
S
Y
C
K
P
S
Y
W
F
A
Z
O
V
D
P
S
E
A
E
P
S
E
A
8
P
S
E
A
7
P
S
E
A
B
P
S
E
A
A
P
S
E
B
N
P
S
E
E
4
P
S
E
P
F
P
S
E
F
S
P
S
E
F
1
P
S
E
K
B
P
S
E
M
N
P
S
E
M
Y
P
S
E
F
5
P
S
E
P
K
P
S
E
P
1
P
S
E
P
B
P
S
E
P
G
P
S
E
P
W
P
S
E
1
4
P
S
E
U
5
P
S
E
U
T
P
S
E
U
2
P
S
E
S
M
F
R
A
N
T
F
R
A
P
2
F
R
A
C
N
F
R
A
C
F
F
R
A
T
N
F
R
A
T
F
F
R
A
T
H
F
R
A
T
O
F
R
A
T
M
F
R
A
T
1
F
R
A
T
E
F
R
A
T
T
F
R
A
T
W
C
Y
C
S
P
H
Y
D
C
U
M
E
T
F
J
M
E
T
N
J
T
H
I
C
A
A
L
I
F
1
A
L
I
F
M
A
L
I
S
L
P
H
O
P
R
V
I
B
A
7
V
I
B
A
E
V
I
B
C
B
V
I
B
C
H
V
I
B
C
J
V
I
B
C
3
V
I
B
C
M
V
I
B
F
N
V
I
B
P
A
V
I
B
T
L
V
I
B
V
U
V
I
B
V
M
V
I
B
V
Y
F
R
A
A
D
L
Y
S
E
N
P
S
E
U
P
P
S
E
U
U
S
T
R
M
K
S
T
R
M
5
X
A
N
A
P
X
A
N
C
8
X
A
N
C
P
X
A
N
C
B
X
A
N
A
C
X
A
N
C
5
X
A
N
O
R
X
A
N
O
M
X
A
N
O
P
X
Y
L
F
A
X
Y
L
F
M
X
Y
L
F
2
X
Y
L
F
T
X
Y
L
F
G
B
A
U
C
H
V
E
S
O
H
R
U
T
M
C
H
A
L
M
S
B
D
E
B
A
D
E
S
B
2
D
E
S
A
A
D
E
S
A
H
D
E
S
T
T
D
E
S
O
H
D
E
S
P
D
D
E
S
S
D
D
E
S
P
S
D
E
S
A
T
D
E
S
R
D
D
E
S
B
D
D
E
S
A
G
D
E
S
D
A
D
E
S
M
R
D
E
S
A
D
D
E
S
V
H
D
E
S
V
M
D
E
S
V
R
D
E
S
V
V
L
A
W
I
P
P
S
E
A
9
P
S
E
P
2
H
I
P
M
A
P
E
L
C
D
P
E
L
P
D
G
E
O
B
B
G
E
O
D
F
G
E
O
L
S
G
E
O
M
G
G
E
O
S
8
G
E
O
S
M
G
E
O
S
L
G
E
O
S
K
G
E
O
U
R
A
N
A
D
2
A
N
A
D
E
A
N
A
D
F
A
N
A
S
K
S
T
I
A
D
C
O
R
C
M
M
Y
X
F
H
M
Y
X
S
D
M
Y
X
X
D
H
A
L
O
1
S
O
R
C
5
D
E
S
A
R
D
E
S
T
A
S
Y
N
A
S
S
Y
N
F
M
A
R
C
B
4
A
R
C
N
C
C
A
M
C
1
C
A
M
C
5
C
A
M
F
F
C
A
M
H
C
C
A
M
J
R
C
A
M
J
D
C
A
M
J
I
C
A
M
J
S
C
A
M
J
M
C
A
M
J
C
C
A
M
J
E
C
A
M
J
J
C
A
M
J
8
C
A
M
L
R
S
U
L
B
S
S
U
L
D
5
H
E
L
A
H
H
E
L
C
P
H
E
L
F
C
H
E
L
H
P
H
E
L
M
1
H
E
L
P
F
H
E
L
P
1
H
E
L
P
5
H
E
L
P
9
H
E
L
P
Y
H
E
L
P
B
H
E
L
P
3
H
E
L
P
C
H
E
L
P
I
H
E
L
P
K
H
E
L
P
L
H
E
L
P
Q
H
E
L
P
G
H
E
L
P
4
H
E
L
P
H
H
E
L
P
7
H
E
L
P
J
H
E
L
P
R
H
E
L
P
2
H
E
L
P
P
H
E
L
P
M
H
E
L
P
S
H
E
L
P
V
S
U
L
K
Y
S
U
L
A
O
S
U
L
D
N
W
O
L
S
U
N
I
T
S
E
N
A
U
P
A
N
I
T
S
B
S
U
L
N
B
B
R
A
H
W
B
R
A
I
P
B
R
A
M
5
B
R
A
P
9
L
E
P
B
P
L
E
P
B
A
L
E
P
B
J
L
E
P
B
L
L
E
P
I
C
L
E
P
I
N
L
E
P
I
I
T
U
R
P
D
B
O
R
C
A
B
O
R
D
L
B
O
R
H
D
B
O
R
R
A
B
O
R
T
9
B
O
R
A
P
B
O
R
B
P
B
O
R
B
U
B
O
R
B
N
B
O
R
B
Z
S
E
D
S
S
S
P
H
C
D
S
P
H
G
B
S
P
H
P
G
S
P
I
A
Z
S
P
I
T
Z
T
R
E
A
Z
T
R
E
B
D
T
R
E
C
H
T
R
E
P
Z
T
R
E
S
6
A
C
E
M
N
A
M
I
C
L
T
H
E
A
S
T
H
E
L
D
M
E
S
F
L
M
Y
C
C
T
M
Y
C
L
9
M
Y
C
L
G
M
Y
C
M
G
M
Y
C
M
S
M
Y
C
P
K
T
H
E
T
1
T
H
E
I
D
T
H
E
G
P
K
O
S
O
T
M
A
R
P
K
P
E
T
M
O
F
E
R
N
B
F
E
R
P
D
T
H
E
A
B
T
H
E
M
4
P
S
E
L
T
T
H
E
M
A
T
H
E
N
R
T
H
E
N
N
T
H
E
P
1
T
H
E
S
Q
M
E
T
I
4
O
P
I
T
P
C
O
R
A
D
A
K
K
M
8
Group 774193ferric uptake regulation protein
Group 774194ferric uptake regulation protein
Group 774195uptake regulation protein
Group 774196Ferric uptake regulator fur family
Group 774197ferric uptake regulation protein
Group 774199Ferric uptake regulator Fur family
Group 774198Ferric uptake regulator Fur family
Group 774202Ferric uptake regulator Fur family
Group 774201Ferric uptake regulator Fur family
Group 774203ferric uptake regulator Fur family
Group 774205ferric uptake regulation protein
Group 774206Ferric uptake regulator Fur family
Group 774208Ferric uptake regulator Fur family
Group 774212Ferric uptake regulator Fur family
Group 774209Ferric uptake regulation protein
Group 774200ferric uptake regulator
Group 774207Ferric uptake regulator Fur family
Group 774211Ferric uptake regulator Fur family
Group 774214regulator Fur family
Group 774213Ferric uptake regulator Fur family
Group 774218uptake regulation protein
Group 774216Ferric uptake regulator Fur family
Group 774219Ferric uptake regulator Fur family
Group 774210Ferric uptake regulator Fur family
Group 774217ferric uptake regulator Fur family COGs COG0735 Fe2+/Zn2+ uptake regulation protein
Group 774204Ferric uptake regulator Fur family
Group 774230Transcriptional regulator Fur family
Group 774220Derived by automated computational analysis using gene prediction method
Group 774221